First wings are tiny buds. Then in some stage a queen pump air into wings (like other insects) and wings get its final size.
At this stage a pupa is white.
Then wings and cuticle dry up.
It varies quite much, when the queen come out from its pupa. Some are quite soft. Swarming queens seems to delay out coming two days and are at once ready to fly.
When wings are in bud stage, they are sensitive to disturbance. So are legs, antennae and cuticle. What ever part of queen may get damages when you shake frames at wrong time. Cutting queen cells produce often damaged queens.