Interestingly in foundationless brood frames, the bees often leave holes for passage between frames, something they can't do in plastic foundation.
Linda, the plastic foundations I use have a hole in each corner for passage to the other side of the frame. When I was looking for the queen on frames with wax foundation, there was a gap at the bottom nearly the length of the frame the queen was on, a lot of bees and I couldn't catch her with such easy access to the other side.
Scooter, sorry, I haven't used foundationless so I can't advise you except to say that often there will be cause to change frames from the brood to the honey super and vice versa, so you will end up with the frames jumbled anyway.
Lone